Network Rail will axe another 500 rail maintenance workers by the end of May through its voluntary redundancy programme.
The cuts follow 235 voluntary redundancies in January.
Network Rail said the redundancies were due to productivity improvements over the last two years.
Maintenance director Steve Featherstone said: “New infrastructure, new technology and new ways of working mean we can maintain the railway more safely and efficiently than ever before with fewer people.
“This is good news for the travelling public. More efficient maintenance means more investment in improving stations, opening new lines and adding capacity to allow more and longer trains.
“It is also good news for our employees – those who are leaving get a severance package and those who remain for the long-term will be part of a flexible and more skilled workforce who can deliver better value for money. That is the best way to safeguard skilled jobs in the future.”